Wow... Ok last try courtesy of a previous post:
The treadmill is acting tangential to the (assumed) free-spinning wheel. A force acting tangential to a free-spinning object can not cause any force on the axis, only a rotational force causing the object to spin about its axis. Therefore the only thing that the treadmill can do is cause the wheels to spin.
The engines on the other hand, are causing a thrusting force which is acting on the wings (or whichever part of the plan that the engine happens to be connected to). Since it is rigidly connected, the force is carried by the member it is connected with towards the CoM (Center of Mass). This transposed force on the Center of Mass causes an acceleration (F=ma). Basic physics principles will explain why and when a force can be translated to a different point. If you read and understand these principles you will note that the force can be translated from its origin to the CoM in a rigid body. Note, also, that on a free-spinning object, a force acting tangent to this object can not be translated to the axle which is the only part of the wheel assembly that is considered to be part of the rigid body of the plane.
Hence, if you use the basic theory of F=ma you will note that the sum of the forces acting on the rigid body of the plane is the thrust and only the thrust. Therefore, the thrust is directly proportional to the acceleration without regard to the force of the treadmill.
Now that was in a "perfect physical world." Very few things change when converting that logic to the real world. These are the following forces that should be taken into account:
Force of friction due to the wheel touching the ground: This can be assumed zero as the tire in both the stated question as well as in real life does not slip relative to the treadmill. If the tire did slip (as if it was locked up) the tire would be considered part of the rigid body and the force of the treadmill would effect the CoM and thus the acceleration. Whether the plane would be able to take off in such a situation would depend on the coefficient of friction between the locked wheel and the ground (very similar to how a seaplane is affected by the water it is in).
Force of friction due to rotation of the wheel about the axle: Most people would agree that a spinning wheel, especially if properly lubricated and fastened as it would be on a plane, causes very little friction on the axle. This friction, however, will add a force that is counteracting the thrust. Once again, though, for this force to cause a plane to not reach take-off speed, the wheel would have to have very high friction, and even then it would still depend on how powerful the engines are (once again, imagine a seaplane and the friction/drag caused by the water, yet it is still able to take off). Therefore, except in extreme rare cases, this will not substantially affect the motion of the plane.
Drag force: The plane is submitted to drag force caused by the air whether it is on a treadmill or not. Airplanes are built with aerodynamics in mind as to minimize this drag force. The only difference might be if the treadmill is causing a velocity in the air, however this would be similar to a breeze in the wind which would cause higher drag forces, but also would move more air over the wings causing higher lift forces. Therefore, in this problem this can also be assumed zero.
Hence, even in a non-perfect physical world, the plane still has a very overwhelming net force moving it forward and will thus fly.
I hope that this clears up any misunderstandings about this problem, and if you made it this far in the post: congratulations. If you made it this far and still don't believe it will fly, go outside and play sports or find something else to do because you are too stubborn to learn physics.